Getting to Know a Grass – Basic Anatomy and Identification

Have you ever tried to identify a grass? Most of us who like to look at plants and learn their names will probably admit that we often give up on grasses pretty quickly, or just ignore them entirely. They aren’t the easiest plants to identify to species, and there are so many of them. Without close inspection, they can all look pretty similar. Their flowers aren’t particularly showy, and their fruits are fairly forgettable. They are strands or clumps of green that create a backdrop for more intriguing forms of vegetation. Yet, they are among the most ecologically and economically important groups of plants on the planet. And actually, if you can ascend the hurdles that come with getting to know them, they are beautiful organisms and really quite amazing.

Kōura in the Grass

The grass family – Poaceae – consists of nearly 8oo genera and about 12,000 species. Grasses occur in a wide range of habitats across the globe. Wherever you are on land, a grass is likely nearby. Grasses play vital roles in their ecosystems and, from a human perspective, are critical to life as we know it. We grow them for food, use them for building materials and fuel, plant them as ornamentals, and rely on them for erosion control, storm water management, and other ecosystem services. We may not acknowledge their presence most of the time, but we very likely wouldn’t be here without them.

The sheer number of grass species is one thing that makes them so difficult to identify. Key identifying features of grasses and grass-like plants (also known as graminoids) tend to be very small and highly modified compared to similar features on other flowering plants. This requires using a hand lens and learning a whole new vocabulary in order to begin to understand a grass’s anatomy. It’s a time commitment that goes beyond a lot of other basic plant identification, and it’s a learning curve that few dare to follow. However, once you learn the basic features, it becomes clear that grasses are relatively simple organisms, and once you start identifying them, it can actually be an exciting and rewarding experience.

Quackgrass (Elymus repens) and Its Rhizome

Depending on the species, grasses can be annuals – completing their life cycle within a single year – or perennials – coming back year after year for two or more years. Most grasses have a fibrous root system; some are quite shallow and simple while others are extremely deep and extensive. Some species of perennial grasses spread by either rhizomes (underground stems), stolons (horizontal, above ground stems), or both. Some grasses also produce tillers, which are essentially daughter plants that form at the base of the plant. The area where roots, rhizomes, stolons, and tillers meet the shoots and leaves of a grass plant is called the crown. This is an important region of the plant, because it allows for regrowth even after the plant has been browsed by a grazing animal or mown down by a lawn mower.

The stem or shoot of a grass is called a culm. Leaves are formed along the lengths of culms, and culms terminate in inflorescences. Leaves originate at swollen sections of the culm called nodes. They start by wrapping around the culm and forming what is called a leaf sheath. Leaves of grasses are generally long and narrow with parallel venation – a trait typical of monocotyledons. The part of the leaf that extends away from the culm is called the leaf blade or lamina. Leaves are alternatively arranged along the length of the stem and are two-ranked, meaning they form two distinct rows opposite of each other along the stem.

The area where the leaf blade meets the leaf sheath on the culm is called the collar. This collar region is important for identifying grasses. With the help of a hand lens, a closer look reveals the way in which the leaf wraps around the culm (is it open or closed?), whether or not there are hairs present and what they are like, if there are auricles (small flaps of leaf tissue at the top of the collar), and what the ligule is like. The ligule is a thin membrane (sometimes a row of hairs) that forms around the culm where the leaf blade and leaf sheath intersect. The size of the ligule and what its margin is like can be very helpful in identifying grasses.

The last leaf on the culm before the inflorescence is called the flag leaf, and the section of the culm between the flag leaf and the inflorescence is called a peduncle. Like the collar, the flower head of a grass is where you’ll find some of the most important features for identification. Grass flowers are tiny and arranged in small groupings called spikelets. In general, several dozen or hundreds of spikelets make up an inflorescence. They can be non-branching and grouped tightly together at the top of the culm, an inflorescence referred to as a spike, or they can extend from the tip of the culm (or rachis) on small branches called pedicels, an inflorescence referred to as a raceme. They can also be multi-branched, which is the most common form of grass inflorescence and is called a panicle.

Either way, you will want to take an even closer look at the individual spikelets. Two small bracts, called glumes, form the base of the spikelet. Above the glumes are a series of florets, which are enclosed in even smaller bracts – the outer bract being the lemma and the inner bract being the palea. Certain features of the glumes, lemmas, and paleas are specific to a species of grass. This includes the way they are shaped, the presence of hairs, their venation, whether or not awns are present and what the awns are like, etc. If the grass species is cleistogamous – like cheatgrass – and the florets never open, you will not get a look at the grass’s sex parts. However, a close inspection of an open floret is always a delight. A group of stamens protrude from their surrounding bracts bearing pollen, while feathery stigmas reach out to collect the pollen that is carried on the wind. Depending on the species, an individual grass floret can have either only stamens, only pistils (the stigma bearing organs), or both. Fertilized florets form fruits. The fruit of a grass is called a caryopsis (with a few exceptions) and is indistinguishable from the seed. This is because the seed coat is fused to the wall of the ovary, unlike other fruit types in which the two are separate and distinct.

If all this doesn’t make you want to run outside and take a close look at some grasses, I don’t know what will. What grasses can you identify in your part of the world? Let me know in the comment section below or check out the linktree and get in touch by the means that suits you best.

Weeds of Boise: Awkward Botany Headquarters

Weeds of Boise: Awkward Botany Headquarters

Last December, Sierra and I left apartment living behind and embarked on a new journey as homeowners, which you can read about in this January’s Year in Review post. This means that Awkward Botany Headquarters now has a yard, and having a yard means we also have weeds.

For many people living in urban areas, the weeds of most concern to them are the ones found in their yards, especially for those that garden or like to keep a tidy yard. Removing weeds is a constant chore. They are always popping up and getting in the way of our plans. In fact, that’s the very definition of a weed – an uninvited plant growing in a location where it isn’t wanted. Despite our best efforts, our yards are always going to have some amount of weeds in them, so what better place to familiarize yourself with your wild urban flora than in your own yard? Or, in this case, our yard.

Our house is located in an area of Boise called the Bench. The Boise Bench, which is actually a series of benches or terraces, is located south of the Boise River and overlooks downtown Boise. The formation of the benches began 2 million years ago as the Boise River cut through the valley. Over time, sediments were deposited at the south bank of the river as it cut further and further northward, leaving behind the series of large terraces. Early in Boise’s history, the Bench was largely agricultural land thanks to the construction of canals. As the city grew, housing and commercial developments expanded across the Bench and have now displaced most of the farmland. Urbanization of the Boise Bench continues today at a steady clip.

While I haven’t had a chance to explore every square inch of the yard, and the growing season is just getting started, what follows are a few photos and a short list of some of the weeds I’ve encountered so far.

  • Arctium minus (burdock)
  • Bromus hordeaceus (soft brome)
  • Bromus tectorum (cheatgrass)
  • Capsella bursa-pastoris (shepherd’s purse)
  • Cichorium intybus (chicory)
  • Cirsium arvense (creeping thistle)
  • Chondrilla juncea (rush skeletonweed)
  • Dactylis glomerata (orchardgrass)
  • Digitaria sanguinalis (crabgrass)
  • Draba verna (spring draba)
  • Echinochloa crus-galli (barnyard grass)
  • Elymus repens (quackgras)
  • Epilobium sp. (willowherb)
  • Erodium cicutarium (redstem filaree)
  • Euphorbia maculata (spotted spurge)
  • Galium aparine (cleavers)
  • Hordeum murinum (wild barley)
  • Lactuca serriola (prickly lettuce)
  • Lepidium sp. (white top)
  • Malva neglecta (common mallow)
  • Matricaria discoidea (pineappleweed)
  • Poa bulbosa (bulbous bluegrass)
  • Polygonum sp. (knotweed)
  • Portulaca oleracea (purslane)
  • Prunella vulgaris (self-heal)
  • Sonchus sp. (sowthistle)
  • Taraxacum officinale (dandelion)
  • Tragopogon dubius (salsify)
  • Ulmus pumila (Siberian elm)
  • Verbena bracteata (bigbract verbena)
  • Veronica sp. (speedwell)

Like all posts in the Weeds of Boise series, this will be updated as I identify and photograph more of the weeds found in this location. Do you have a yard in an urban area? What weeds are you seeing in your yard this year? Let us know in the comment section below.